Review of the Gymnopleurini (Coleoptera: Scarabaeidae: Scarabaeinae). II. The genus Paragymnopleurus Shipp

Abstract

The genus Paragymnopleurus Shipp, 1897 (Coleoptera: Scarabaeidae: Scarabaeinae: Gymnopleurini) is
characterized and its constituent taxa are keyed and illustrated. Twelve species and five subspecies are deemed valid,
and five species groups are recognized. Three new synonymies include: Paragymnopleurus stipes japonicus Balthasar
is synonymized with P. ambiguus Janssens, and P. maurus malayanus Ochi and Kon and P. maurus pauliani Janssens
are synonymized with the nominotypical subspecies. First country and provincial records are reported for P. brahminus
(Waterhouse), P. maurus (Sharp) and P. sinuatus szechouanicus Balthasar. A lectotype is here designated for
Gymnopleurus singularis Waterhouse, validating an unpublished designation. A checklist of valid species and synonyms
is provided.